About Me

I am a Ph.D. candidate in Biological Anthropology working with Dr. Evan MacLean. I graduated from the University of North Carolina at Chapel Hill in 2016 with a B.S. in Psychology and minor in Anthropology, and I received an M.A. in Anthropology from the University of Arizona in 2018.

Prior to joining the School of Anthropology at the University of Arizona, I worked in a number of diverse research roles, including projects examining human pain perception, metacognition, and memory at UNC, user experience (UX) at Lenovo, and canine and non-human primate cognition in Yale University’s Canine Cognition Center and with Yale’s Comparative Cognition Laboratory at the Cayo Santiago Biological Field Station in Puerto Rico.

My research interests broadly center on cognitive evolution, with a particular focus on sociocognitive abilities relating to theory of mind and shared intentionality. I am also interested in relationships between neuroanatomy and cognition both within and across species. I investigate these topics by integrating non-invasive cognitive, behavioral, and biological research in dogs, wolves, and free-ranging non-human primates.

I am currently affiliated with the Cognitive Science Graduate Interdisciplinary Program, and have worked as a Graduate Teaching Associate in both the School of Anthropology and the School of Mind, Brain and Behavior at the University of Arizona. In the summer of 2021, I will be joining the Department of Psychology at Yale University as a postdoctoral associate.

Research Experience

Wildlife Science Center, Stacy, MN (2018 – Present)
Humane Society of Southern Arizona, Tucson, AZ (2018 – Present)
Arizona Canine Cognition Center, Tucson, AZ (2016 – Present)
Canine Companions for Independence, Santa Rosa, CA & Oceanside, CA (2016 – Present)
Cayo Santiago Biological Field Station, Punta Santiago, Puerto Rico (2016 – Present)
Canine Cognition Center at Yale, New Haven, CT (2015)
Human Memory Lab, Chapel Hill, NC (2015 – 2016)
Lenovo User Experience Research Team, Morrisville, NC (2014 – 2016)
Somatosensory Research Lab, Chapel Hill, NC (2013 – 2014)

Peer-Reviewed Publications

Bray, E.E., Gruen, M.E., Gnanadesikan, G.E., Horschler, D.J., Levy, K.M., Kennedy, B.S., Hare, B.A., & MacLean, E.L. (2020). Dog cognitive development: A longitudinal study across the first two years of life. Animal Cognition. [PDF]

Horschler, D.J., MacLean, E.L., & Santos, L.R. (2020). Advancing gaze-based research on primate theory of mind. Trends in Cognitive Sciences, 24(10), 778-779. [PDF]

Bray, E.E., Gruen, M.E., Gnanadesikan, G.E., Horschler, D.J., Levy, K.M., Kennedy, B.S., Hare, B.A., & MacLean, E.L. (2020). Cognitive characteristics of 8-to-10-week-old assistance dog puppies. Animal Behaviour, 66, 193-206. [PDF]

Horschler, D.J., MacLean, E.L., & Santos, L.R. (2020). Do non-human primates really represent others’ beliefs? Trends in Cognitive Sciences, 24(8), 594-605. [PDF]

Horschler, D.J. & MacLean, E.L. (2019). Leveraging brain-body scaling relationships for comparative studies. Animal Cognition, 22(6), 1197-1202. [PDF]

Horschler, D.J., Santos, L.R., & MacLean, E.L. (2019). Do non-human primates really represent others' ignorance? A test of the awareness relations hypothesis. Cognition, 190, 72-80. [PDF]

Horschler, D.J., Hare, B., Call, J., Kaminski, J., Miklósi, Á., & MacLean, E.L. (2019). Absolute brain size predicts dog breed differences in executive function. Animal Cognition, 22(2), 187-198. [PDF]
    Selected as the issue’s highlight article

Submitted Manuscripts

Arre, A.M.* & Horschler, D.J.* (In revision). Swimming and diving as social play in juvenile rhesus macaques (Macaca mulatta). Behaviour. [*Denotes equal contribution]

Horschler, D.J., Santos, L.R., & MacLean, E.L. (Under review). How do non-human primates represent others’ awareness of hidden objects? Cognition.

Bray, E.E., Gnanadesikan, G.E., Horschler, D.J., Levy, K.M., Kennedy, B.S., Famula, T.R. & MacLean, E.L. (Under review). Early-emerging and highly-heritable sensitivity to human communication in dogs. Science.

Manuscripts in Preparation

Horschler, D.J., Bray, E.E., Gnanadesikan, G.E., Byrne, M., & MacLean, E.L. (In prep). Dogs re-engage human partners when joint social play is interrupted.

Salomons, H., Smith, K., Callahan-Beckel, M., Callahan, M., Levy, K., Kennedy, B., Gnanadesikan, G.E., Horschler, D.J., Tan, J., MacLean, E.L., & Hare, B. (In prep). Early-emerging social cognition in dogs but not wolves.

Conference Presentations

Horschler, D.J., Bray, E.E., & MacLean, E.L. (2020). Do dogs re-engage humans when joint experiences end? Talk presented at the Yale University East Coast Canine Cognition Workshop, New Haven, CT.

Bray, E.E., Gnandesikan, G.E., Horschler, D.J., & MacLean, E.L. (2020). Early development and longitudinal stability of cognitive traits in working dogs. Talk presented at the Yale University East Coast Canine Cognition Workshop, New Haven, CT.

MacLean, E.L., Bray, E.E., Gnanadesikan, G.E., & Horschler, D.J. (2020). Associations between individual differences in cognition and training outcomes in assistance dogs. Talk presented at the Yale University East Coast Canine Cognition Workshop, New Haven, CT.

Horschler, D.J., Santos, L.R., & MacLean, E.L. (2019). Awareness relations underpin knowledge representation in rhesus monkeys. Talk presented at the UA-ASU Cognitive Science Conclave, Tucson, AZ.

Horschler, D.J., Santos, L.R., & MacLean, E.L. (2019). How do non-human primates represent what others know? Talk presented at the 7th Annual Conference of the Southwestern Association of Biological Anthropologists, Tempe, AZ.
    Awarded prize for best student podium presentation

MacLean, E.L., Bray, E.E., Gnanadesikan, G.E., & Horschler, D.J. (2019). Ontogeny and heritability of cognitive and temperamental traits in an assistance dog population. Talk presented at the International Canine Science Conference, Tempe, AZ.

Horschler, D.J., Santos, L.R., & MacLean, E.L. (2019). Do non-human primates really represent others' ignorance? Talk presented at the 26th International Conference on Comparative Cognition, Melbourne Beach, FL.
    Selected as a finalist for the Ron Weisman Outstanding Student Presentation Award

Bray, E.E., Gnanadesikan, G.E., Horschler, D.J., & MacLean, E.L. (2019). Early emerging cognition in 9-week-old puppies. Talk presented at the 26th International Conference on Comparative Cognition, Melbourne Beach, FL.

MacLean, E.L., Bray, E.E., Gnanadesikan, G.E., & Horschler, D.J. (2019). Heritability of cognitive traits in a pedigreed dog population. Talk presented at the 26th International Conference on Comparative Cognition, Melbourne Beach, FL.

Horschler, D.J. & MacLean, E.L. (2018). Dogs as a model for the cognitive implications of neuroanatomical variation. Talk presented at the Yale University East Coast Canine Cognition Workshop, New Haven, CT.

MacLean, E.L. & Horschler, D.J. (2018). Breed differences in dog cognition are predicted by absolute brain volume. Talk presented at the 126th Annual Convention of the American Psychological Association, San Francisco, CA.

Horschler, D.J. & MacLean, E.L. (2017). Absolute brain size predicts dog breed differences in executive function and social cognition. Poster presented at the UA-ASU Cognitive Science Conclave, Tucson, AZ.

Horschler, D.J. & MacLean, E.L. (2017). Absolute brain size predicts dog breed differences in working memory and social cognition. Poster presented at the North American Canine Science Forum, Tempe, AZ.

Internal Presentations

Horschler, D.J. (2020). How is human social cognition different? Invited talk presented via Zoom for PHIL 271: Philosophy of Language at Yale University.

Horschler, D.J., Santos, L.R., & MacLean, E.L. (2019). Do monkeys understand knowledge and ignorance? Talk presented at the Anthropology Graduate Students at the University of Arizona (AGUA) Student Lecture Series, Tucson, AZ.

Horschler, D.J. (2019). What can dogs teach us about cognitive evolution? Talk presented at the UA Grad Slam Three-Minute Thesis Competition, Tucson, AZ.
    Selected as a semifinalist in campus-wide competition

Gnanadesikan, G.E., Horschler, D.J., & MacLean, E.L. (2019). Social cues and hormonal profiles over development in wolf puppies. Poster presented at the GPSC Student Showcase, Tucson, AZ.

Horschler, D.J. & MacLean, E.L. (2018). Absolute brain size predicts dog breed differences in executive function and social cognition. Poster presented at the GPSC Student Showcase, Tucson, AZ.

Horschler, D.J., Susser, J.A., & Mulligan, N.W. (2016). Metamemory for performed and observed actions: Within-subjects. Poster presented at the UNC Psychology and Neuroscience Honors Poster Symposium, Chapel Hill, NC.

Horschler, D.J., Susser, J.A., & Mulligan, N.W. (2016). Metamemory for performed and observed actions: Within-subjects. Poster presented at the UNC Celebration of Undergraduate Research, Chapel Hill, NC.

Horschler, D.J. (2015) A brief overview of canine cognition research. Invited talk presented for PSYC 67: The Senses of Animals at the University of North Carolina, Chapel Hill, NC.

Horschler, D.J. (2014). Insights into user experience at Lenovo. Poster presented at the Karen M. Gil Internship Program Poster Symposium, Chapel Hill, NC.

Selected Media Coverage

StarTribune

What separates dogs and wolves? Researchers journey to Anoka County to find out

Coverage of Salomons, H., Smith, K., Callahan-Beckel, M., Callahan, M., Levy, K., Kennedy, B., Gnanadesikan, G.E., Horschler, D.J., Tan, J., MacLean, E.L., & Hare, B. (In prep). Early-emerging social cognition in dogs but not wolves.


Commentary on Related Research

Teaching Experience

NSCS 200: Fundamentals of Neuroscience and Cognitive Science (Spring 2020)
Department of Neuroscience | School of Mind, Brain & Behavior | College of Science
Graduate Teaching Associate | ~160 students

ANTH 170C2: Animal Minds (Fall 2019)
School of Anthropology | College of Social and Behavioral Sciences
Graduate Teaching Associate | Two sections (~30 students each)

ANTH 327: Dog Thought (Spring 2019)
School of Anthropology | College of Social and Behavioral Sciences
Graduate Teaching Associate | ~150 students

NSCS 321: Research Methods in Cognitive Science (Fall 2018)
Department of Neuroscience | School of Mind, Brain & Behavior | College of Science
Graduate Teaching Assistant | ~110 students

ANTH 170: Human Variation in the Modern World (Fall 2017 & Spring 2018)
School of Anthropology | College of Social and Behavioral Sciences
Graduate Teaching Assistant | Two sections per semester (~30 students each)

Contact Information

Mailing Address
The University of Arizona
School of Anthropology
P.O. Box 210030
Tucson, AZ 85721

Email Address
horschler [at] email.arizona.edu

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